Psyllipsocetae
Psyllipsocidae
Emilie Bess and Kevin P. JohnsonThis tree diagram shows the relationships between several groups of organisms.
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close boxIntroduction
The family Psyllipsocidae includes about 50 species in 5 genera distributed worldwide. In North America, six species in four genera are known.Psyllipsocids live in caves, on rock surfaces, in dead leaves, and in buildings. Five of the six North American species have been collected in homes, office buildings, or cellars. These are small bark lice and their bodies may be hairy but do not have scales.
Characteristics
Synapomorphy:
In the female. the spremathecal sac has complex sclerifications at the origin and usually has an accessory vesicle.
General Characters:
Head: Head is long due to long cheeks (gena).
Antennae are long and have more than 20 segments without ringed sculpturing (annulations).
Ocelli are not in groups.
Legs: Tarsi have 3 segments.
Wings: Wings are sometimes hairy but do not have scales.
Forewing: Pterostigma is not thickened.
Vein Sc has a short basal segment that ends at or before wing margin.
Veins CuP and A1 meet on the wing margin forming nodulus.
Hindwing vein A is not branched or has two branches.
Male: Paraproct has an anal spine.
Phallosome is composed of two curved arms that are never fused anteriorly.
Female: Paraproct has an anal spine.
Gonapophyses are reduced:
Dorsal valve is small or absent.
Ventral valve is very small.
External valve is broad and hairy.
Eggs are laid singly and are not covered with silk or debris.
Discussion of Phylogenetic Relationships
The monophyly of family Psyllipsocidae is supported by one character of the female genitalia: the spremathecal sac with complex sclerifications at the origin and usually with an accessory vesicle. Molecular analysis that included the genera Psyllipsocus and Dorypteryx also supports monophyly of the family (18S & Histone3 nDNA, 16S rDNA; Yoshizawa et al. 2006).References
Lienhard, C. and C.N. Smithers. 2002. Psocoptera (Insecta) World Catalogue and Bibliography. Muséum d'Histoire Naturelle, Geneva, Switzerland.
Mockford, E.L. 1993. North American Psocoptera (Insecta). Gainesville, Florida: Sandhill Crane Press.
Smithers, C.N. 1996. Psocoptera. Pp. 1-80, 363-372 (Index) in Wells A. (ed.) Zoological Catalogue of Australia. Vol. 26. Psocoptera, Phthiraptera, Thysanoptera. Melbourne: CSIRO Publishing, Australia.
Smithers, C.N. 1972. The classification and phylogeny of the Psocoptera. Memoirs of the Australian Museum 14: 1–349.
Yoshizawa, K., C. Lienhard, and K.P. Johnson. 2006. Molecular systematics of the suborder Trogiomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 146: 287–299.
About This Page
Emilie Bess
Illinois Natural History Survey, Champaign, Illinois, USA
Kevin P. Johnson
Illinois Natural History Survey, Champaign, Illinois, USA
Correspondence regarding this page should be directed to Emilie Bess at and Kevin P. Johnson at
Page copyright © 2009 Emilie Bess and Kevin P. Johnson
All Rights Reserved.
- First online 18 March 2009
- Content changed 18 March 2009
Citing this page:
Bess, Emilie and Kevin P. Johnson. 2009. Psyllipsocetae. Psyllipsocidae. Version 18 March 2009 (temporary). http://tolweb.org/Psyllipsocetae/14453/2009.03.18 in The Tree of Life Web Project, http://tolweb.org/